International Journal of Horticulture, 2025, Vol.15, No.1, 8-20 http://hortherbpublisher.com/index.php/ijh 19 Funding This study was supported by Hainan Province Science and Technology Special Fund (Grant No: ZDYF2021XDNY120). Conflict of Interest Disclosure The authors affirm that this research was conducted without any commercial or financial relationships that could be construed as a potential conflict of interest. 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