Plant Gene and Trait 2025, Vol.16, No.1, 1-14 http://genbreedpublisher.com/index.php/pgt 12 Funding This study was supported by the Research and Training Fund of the Hainan Institute of Tropical Agricultural Resources (Project No. H2025-01). Conflict of Interest Disclosure The authors affirm that this research was conducted without any commercial or financial relationships that could be construed as a potential conflict of interest. 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