International Journal of Molecular Evolution and Biodiversity, 2025, Vol.15, No.2, 84-98 http://ecoevopublisher.com/index.php/ijmeb 96 In addition, root traits show irreplaceable importance in drought adaptation. Varieties with deep roots and well-developed root branches have stronger water and mineral nutrient absorption capacity, which helps to alleviate the adverse effects of drought on plant growth. Future breeding strategies should fully integrate the traits of root development regulation and water use efficiency to improve resource adaptability. It is worth noting that biological control agents such as Trichoderma harzianum have shown application potential in enhancing plant drought resistance. By affecting plant hormone balance and rhizosphere microecological system, they introduce eco-friendly intervention methods for sustainable agricultural systems. Acknowledgments Thank you very much to the reviewers for providing detailed comments and guidance on the manuscript of this study. The reviewers' keen insights into the issues and attention to detail have greatly benefited the author. Conflict of Interest Disclosure The author affirms that this research was conducted without any commercial or financial relationships that could be construed as a potential conflict of interest. References Araus J., Slafer G., Reynolds M., and Royo C., 2002, Plant breeding and drought in C3 cereals: what should we breed for? Annals of botany, 89: 925-940. https://doi.org/10.1093/AOB/MCF049 Arroyo-Álvarez E., Chan-León A., Girón-Ramírez A., Fuentes G., Estrella-Maldonado H., and Santamaría J., 2023, Genome-wide analysis of WRKY and NAC transcription factors in Carica papaya L. and their possible role in the loss of drought tolerance by recent cultivars through the domestication of their wild ancestors, Plants, 12(15): 2775. https://doi.org/10.3390/plants12152775 Ashraf M., 2010, Inducing drought tolerance in plants: recent advances, Biotechnology advances, 28(1): 169-183. https://doi.org/10.1016/j.biotechadv.2009.11.005 Batool M., El-Badri A., Wang Z., Mohamed I., Yang H., Ai X., Salah A., Hassan M., Sami R., Kuai J., Wang B., and Zhou G., 2022, Rapeseed morpho-physio-biochemical responses to drought stress induced by PEG-6000, Agronomy, 12(3): 579. https://doi.org/10.3390/agronomy12030579 Begna T., 2022, Breeding strategies for improvement of drought tolerant in crops, International Journal of Research in Agronomy, 18(3): 177-184. https://doi.org/10.33545/2618060x.2022.v5.i1a.99 Bhuiyan T., Ahamed K., Nahar K., Mahmud J., Bhuyan M., Anee T., Fujita M., and Hasanuzzaman M., 2019, Mitigation of PEG-induced drought stress in rapeseed (Brassica rapa L.) by exogenous application of osmolytes, Biocatalysis and Agricultural Biotechnology, 20: 101197. https://doi.org/10.1016/J.BCAB.2019.101197 Bouchereau A., Clossais-Besnard N., Bensaoud A., Leport L., and Renard M., 1996, Water stress effects on rapeseed quality, European Journal of Agronomy, 5: 19-30. https://doi.org/10.1016/S1161-0301(96)02005-9 Cao B., Bai J., Wang X., Zhang Y., Yu X., Hu S., and He Y., 2022, BnA.JAZ5 attenuates drought tolerance in rapeseed through mediation of ABA-JA Crosstalk, Horticulturae, 8(2): 131. https://doi.org/10.3390/horticulturae8020131 Cattivelli L., Rizza F., Badeck F., Mazzucotelli E., Mastrangelo A., Francia E., Marè C., Tondelli A., and Stanca A., 2008, Drought tolerance improvement in crop plants: An integrated view from breeding to genomics, Field Crops Research, 105: 1-14. https://doi.org/10.1016/J.FCR.2007.07.004 Chaghakaboodi Z., Kakaei M., and Zebarjadi A., 2021, Study of relationship between some agro-physiological traits with drought tolerance in rapeseed (Brassica napus L.) genotypes, Central Asian Journal of Plant Science Innovation, 1(1): 1-9. https://doi.org/10.22034/CAJPSI.2021.01.01 Channaoui S., Idrissi I., Mazouz H., and Nabloussi A., 2019, Reaction of some rapeseed (Brassica napus L.) genotypes to different drought stress levels during germination and seedling growth stages, OCL, 26: 23. https://doi.org/10.1051/OCL/2019020 Dai L., Li J., Harmens H., Zheng X., and Zhang C., 2020, Melatonin enhances drought resistance by regulating leaf stomatal behaviour, root growth and catalase activity in two contrasting rapeseed (Brassica napus L.) genotypes, Plant Physiology and Biochemistry: PPB, 149: 86-95. https://doi.org/10.1016/j.plaphy.2020.01.039 Germchi S., Shekari F., Hassanpooraghdam M., Khorshidi M., and Shekari F., 2010, Water deficit stress affects growth and some biochemical characteristics of rapeseed (Brassica napus L.), Journal of Food Agriculture and Environment, 8: 1126-1129. Jiang Y., Liang G., and Yu D., 2012, Activated expression of WRKY57 confers drought tolerance in Arabidopsis, Molecular Plant, 5(6):1375-1388. https://doi.org/10.1093/mp/sss080
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